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Soldier beetle

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Soldier beetle
Temporal range: Albian–Present
Chauliognathus lugubris.jpg
Chauliognathus lugubris
Scientific classification e
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Coleoptera
Suborder: Polyphaga
Infraorder: Elateriformia
Superfamily: Elateroidea
Family: Cantharidae
Imhoff, 1856
Synonyms

Chauliognathidae

The soldier beetles (Cantharidae) are relatively soft-bodied, straight-sided beetles. They are cosmopolitan in distribution. One of the first described species has a color pattern reminiscent of the red coats of early British soldiers, hence the common name. They are also known commonly as leatherwings because of their soft elytra.[1]

Historically, these beetles were placed in a superfamily "Cantharoidea", which has been subsumed by the superfamily Elateroidea; the name is still sometimes used as a rankless grouping, including the families Cantharidae, Lampyridae, Lycidae, Omethidae (which includes Telegeusidae), Phengodidae, and Rhagophthalmidae.

Evolutionary history

The oldest described member of the family is Molliberus from the early Albian aged El Soplao amber from Cantabria, Spain, belonging to the tribe Cantharini in the subfamily Cantharinae. Other described genera include 6 from the Cenomanian aged Burmese amber, with 5 belonging to Cantharinae and one to Malthininae, and Katyacantharis, from the Cenomanian aged Agdzhakend amber of Azerbaijan, suggested to belong to Cantharinae. Indeterminate specimens have been reported from the Aptian Koonwarra fossil bed of the Strzelecki Group, Australia and the Barremian aged Lebanese amber.[2][3]

Subfamilies, tribes and selected genera

Five subfamilies are normally accepted:

Cantharinae

  • tribe Cantharini
  • tribe Podabrini
    • Podabrus

Chauliognathinae

  • tribe Chauliognathini
    • Belotus
    • Chauliognathus
  • tribe Ichtyurini
    • Trypherus

Dysmorphocerinae

  • Afronycha Wittmer, 1949 - central-southern Africa
  • Asilis Broun, 1893 - New Zealand
  • Compsonycha
  • Dysmorphocerus Solier, 1849
  • Flabelloontelus
  • Geigyella Wittmer, 1972 - New Guinea
  • Hansasilis
  • Heteromastix Boheman, 1858 - Australia
  • Hyponotum
  • Micronotum
  • Neoontelus Wittmer, 1972 - New Zealand
  • Oontelus Solier, 1849 - S. America
  • Plectocephalon
  • Plectonotum Gorham, 1891 - Americas

Malthininae

  • tribe Malchinini
    • Macrocerus Motschulsky, 1845 - Europe (synonym Malchinus)
  • tribe Malthinini
    • Caccodes Sharp, 1885 - central America, Pacific islands
    • Malthinellus Kiesenwetter, 1874 - Japan
    • Malthinus Latreille, 1805 - Japan, Europe, N. America
  • tribe Malthodini
    • Frostia Bert. ex Guill.
    • Malthodes Kiesenwetter, 1852 - mostly Europe, N. America & New Zealand
    • Archaeomalthodes Hsiao et al. 2016 Burmese amber, Myanmar, Cenomanian

Silinae

  • tribe Silini
    • Cordylocera Guérin-Méneville, 1830
    • Silis Charpentier, 1825
  • tribe Tytthonyxini
    • Tytthonyx LeConte, 1851

Reproduction

Large males of the soldier beetle exercise choice for larger females. Body size correlates with the abilities of males to secure females, and of females to evade males.[4]

See also

References

  1. ^ Phillips, C., et al. Leatherwing (Soldier) Beetles. Virginia Cooperative Extension, Virginia Tech and Virginia State University. 2013.
  2. ^ Peris, David; Fanti, Fabrizio (November 2018). "Molliberus albae gen. et sp. nov., the oldest Laurasian soldier beetle (Coleoptera: Cantharidae) from the Lower Cretaceous Spanish amber". Cretaceous Research. 91: 263–268. doi:10.1016/j.cretres.2018.07.003.
  3. ^ Kazantsev, Sergey V.; Perkovsky, Evgeny E. (2019-02-28). "The first Cretaceous beetle from Azerbaijan: Katyacantharis zherikhini gen. et sp. n. (Coleoptera, Cantharidae) from Cenomanian Agdzhakend amber". Palaeoentomology. 2 (1): 7. doi:10.11646/palaeoentomology.2.1.2. ISSN 2624-2834.
  4. ^ McLain, Denson K.; Pratt, Ann E.; Shure, Donald J. (June 2015). "Size dependence of courtship effort may promote male choice and strong assortative mating in soldier beetles". Behavioral Ecology and Sociobiology. 69 (6): 883–894. doi:10.1007/s00265-015-1900-6. ISSN 0340-5443.

External links

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